I followed the link as well as I could, and I'm not sure I made it to where you were pointing.
As near as I can tell you were pointing out winter kill, and human predation as factors in the decline of elk in the northern yellowstone.
Yes there have been bad winter kills, a friend sent me a clipping years ago (pre-internet) probably still here somewhere. The gist of which was the winter kill was so bad the predators/scavengers couldn't keep up. The clean up crew of that time were generally a bit lazier, and tended to finish a higher per cent of an animal before killing another.
Human predation can be regulated.
The reproductive rate of ungulates can produce quick recoveries, in these scenario's as neither really impacted the calf crop like the wolves have. I think it was the count of 2013 that failed to come up with a calf that made it into the next year. Single digit survival has been noted in several places.
Winter, and human hunting have seasons, and limits. Attempts to place limits on wolves is the issue. Recreational hunting/trapping/SSS/auto interactions to this point have failed to maintain wolves at any particular level.
Neither can it be denied (although they try) the wolves impact on other predators, grizzly dispersal from Yellowstone, single digit mountain lion production, etc. Or the contribution to the complete removal of caribou from the lower 48.
The current model of fair chase will have to be modified significantly if we are to manage this predator in a somewhat cost effective way. Guess what? wolves shot from helicopters on our dime are just as dead, as if the fly overs were conducted privately.