Folks not believing nor understanding how a rimless bottleneck case fits the front of the chamber perfectly centered might consider a few things that explains why and how.
First, inspect your bolt action rifle and notice the one or two things that push the cartridge forward into the chamber until it stops after the bolt's closed.
Second, figure out why the case shoulder being forced into the chamber shoulder will center it there. Note the two angles, both shoulder and chamber, are the same for all practical purposes. Such cases do this the same way headspace gauges do and these gauges do center perfectly in the chamber's shoulder when all the way forward without any neck at all.
Third, if you cannot figure out the first two, then measure the clearance around a case neck to the chamber neck when the round's chambered and pushed all the way forward by those one or two external forced putting it there. There's an easy way to do this; cut the barrel off just in front of the chamber mouth, square it up then use a 10X loupe with a scale in thousandths to do the measuring.
Measure the diameter of a resized case neck (to the nearest .0001"), then compare that to the measured diameter of the chamber neck at the same point, you may be surprised to see how much clearance there is. If there's clearance, then there's no way that area on both the case and in the chamber will do anything to center the case neck in the chamber neck.
Then you may be able to understand why a .243 Win. cartridge will fit a .308 Win. chamber such that when fired, its case neck and therefore the bullet, too, is perfectly centered in the bore. Even if its body diameters are .006" smaller than the chamber and there's .006" head clearance due the case headspace being .006" shorter than chamber headspace.
Some folks have been sizing fired cases this way for decades and getting accuracy equal to or better than what the benchresters do. Sierra Bullets for one; their best match bullets go into sub 1/4 MOA groups in their 200 yard indoor range in Missouri. I've seen several 10-shot test groups from their 100 yard range in California testing 168's in .308 Win. cases that were in the ones; under 2/10ths inch.
First, inspect your bolt action rifle and notice the one or two things that push the cartridge forward into the chamber until it stops after the bolt's closed.
Second, figure out why the case shoulder being forced into the chamber shoulder will center it there. Note the two angles, both shoulder and chamber, are the same for all practical purposes. Such cases do this the same way headspace gauges do and these gauges do center perfectly in the chamber's shoulder when all the way forward without any neck at all.
Third, if you cannot figure out the first two, then measure the clearance around a case neck to the chamber neck when the round's chambered and pushed all the way forward by those one or two external forced putting it there. There's an easy way to do this; cut the barrel off just in front of the chamber mouth, square it up then use a 10X loupe with a scale in thousandths to do the measuring.
Measure the diameter of a resized case neck (to the nearest .0001"), then compare that to the measured diameter of the chamber neck at the same point, you may be surprised to see how much clearance there is. If there's clearance, then there's no way that area on both the case and in the chamber will do anything to center the case neck in the chamber neck.
Then you may be able to understand why a .243 Win. cartridge will fit a .308 Win. chamber such that when fired, its case neck and therefore the bullet, too, is perfectly centered in the bore. Even if its body diameters are .006" smaller than the chamber and there's .006" head clearance due the case headspace being .006" shorter than chamber headspace.
Some folks have been sizing fired cases this way for decades and getting accuracy equal to or better than what the benchresters do. Sierra Bullets for one; their best match bullets go into sub 1/4 MOA groups in their 200 yard indoor range in Missouri. I've seen several 10-shot test groups from their 100 yard range in California testing 168's in .308 Win. cases that were in the ones; under 2/10ths inch.